Year of Publication 2023, Vol. 75 (2)

Date Published 15 November 2023
D.J. Middleton

Flora of Singapore precursors 40: Notes on Ericaceae in Singapore [Page 177 - 181]
The nomenclature and typification of the species of Ericaceae in Singapore are discussed. A neotype is designated for Leucopogon malayanus Jack. Lectotypes are designated for Rhododendron longiflorum Lindl., R. javanicum (Blume) Benn. var. tubiflorum Hook.f., Vaccinium bracteatum Thunb. and, in a second step, V. malaccense Wight.
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D.M. Johnson & N.A. Murray

A contribution to the systematics of Xylopia (Annonaceae) in the New Guinea region [Page 183 - 255]
In the New Guinea region, including the Solomon Islands and Halmahera, three species of the pantropical genus Xylopia L. have been previously recognised: X. calosericea Diels, X. papuana Diels, and X. peekelii Diels. In recent floristic tabulations the genus has thus comprised a minor component of the Annonaceae flora. Here we present descriptions and illustrations of 14 new species of the genus from the region: Xylopia aenea D.M.Johnson & N.A.Murray, X. ampla D.M.Johnson & N.A.Murray, X. brunneola D.M.Johnson & N.A.Murray, X. bullata D.M.Johnson & N.A.Murray, X. chlorosperma D.M.Johnson & N.A.Murray, X. cornuta D.M.Johnson & N.A.Murray, X. corrugata D.M.Johnson & N.A.Murray, X. makiraensis D.M.Johnson & N.A.Murray, X. musella D.M.Johnson & N.A.Murray, X. pachysericea D.M.Johnson & N.A.Murray, X. rogstadii D.M.Johnson & N.A.Murray, X. sulangwane D.M.Johnson & N.A.Murray, X. takeuchii D.M.Johnson & N.A.Murray, and X. vulcanicola D.M.Johnson & N.A.Murray. We distinguish the 17 species in a diagnostic key. The region includes Xylopia species that are among the tallest in this genus of c. 220 species, one with the largest leaves and another with the largest fruit, as well as several species with distinctive cataphyll-covered resting buds. The genus has undergone multiple dispersals both to and within the region, with higher diversity on smaller islands than on New Guinea. Its presence on oceanic islands indicates over-water dispersal.
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Z. Ezedin

Confirmation of Morinda bracteata (Rubiaceae) in New Guinea [Page 257 - 262]
The tree species Morinda bracteata Roxb. is here confirmed for New Guinea after having been previously overlooked. A brief summary of its taxonomic history is given, along with its revised distribution which now appears restricted to the east of Huxley’s Line when Palawan is included.
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D.J. Middleton

A revision of Codonoboea (Gesneriaceae: Didymocarpoideae) in Thailand [Page 263 - 296]
The genus Codonoboea Ridl. (Gesneriaceae: Didymocarpoideae: Trichosporeae: Didymocarpinae) is revised for Thailand. Thirteen species are recognised, one of which, Codonoboea poopathii D.J.Middleton, is new to science and three of which, C. dawnii (Kiew) Kiew, C. oreophila Kiew ex C.L.Lim and C. urticoides (A.Weber) Kiew, are new records for Thailand. Didymocarpus reptans Jack is neotypified; D. hispidus Ridl. var. selangorensis Ridl., D. inaequalis Ridl., D. rugosus Ridl. and D. urticifolius Ridl. are lectotypified; and D. hispidus Ridl. is lectotypified in a second step. A key to the species is provided and all species are described.
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D.J. Middleton

Four new species of Gesneriaceae from Vietnam [Page 297 - 307]
The new species Allocheilos villosus B.L.Burtt ex D.J.Middleton, Didymocarpus tamdaoensis D.J.Middleton, Didymocarpus dalatensis D.J.Middleton and Loxostigma vietnamensis D.J.Middleton from Vietnam are described.
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Middleton, D.J.

A new species of Paraboea (Gesneriaceae: Didymocarpoideae) from Myanmar [Page 309 - 314]
The new species Paraboea babae D.J.Middleton from Shan State in Myanmar is described and illustrated.
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W.W. Seah

A name change in Gynochthodes (Rubiaceae) [Page 319 - 320]
A new combination Gynochthodes wongiana (Suratman) W.W.Seah is proposed here based on Morinda wongiana Suratman and the name Gynochthodes suratmanii K.M.Wong & Razafim. is placed in its synonymy.
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Year of Publication 2023, Vol. 75 (1)

Date Published 12 June 2023
Er, K.B.H., Chong, K.Y., Choo, T.Y.S., Middleton, D.J. & Tan, P.Y.

Establishing a network of long-term forest monitoring plots in Singapore [Page 1 - 20]
We review the history of plot-based studies of forests in Singapore and their contribution to our understanding of tropical forest ecology, especially of the regenerative capabilities of forest remnants after fragmentation, land-use change, and other disturbances. With this, we describe the establishment of the Long-Term Forest Ecological Monitoring plot network that includes the continued utilisation of sets of recently established, standardised plots along with the re-establishment of a historical set of plots surveyed by a team led by Wong Yew Kwan in 1992, ensuring the plot design is harmonised across the network.
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Niissalo, M.A., Leong, P.K.F., Tay, F.E.L., Choo, L.M., Kurzweil, H. & Khew, G.S.

A new species of Claderia (Orchidaceae) [Page 21 - 41]
A little-studied orchid genus, Claderia Hook.f., has until now contained just one or two species, Claderia viridiflora Hook.f. and its possible synonym, C. papuana Schltr. We describe a new species from Singapore, which differs from these Claderia in having small, cream-yellow flowers that are nodding and presented in a long, unbranched terminal inflorescence, mostly with two flowers open simultaneously. We name the new species Claderia leontocampus Niissalo. Plants of the new species have been collected or recorded in Sumatra, Peninsular Malaysia and possibly Peninsular Thailand. We carried out population genetics analyses to show distinction between the new species and Claderia viridiflora. The two species occur in the same habitat in Singapore. We used phylogenetic analyses to find out the phylogenetic position of Claderia within Orchidaceae; the genus is here considered to belong to subfamily Epidendroideae Kostel., tribe Epidendreae Lindl., subtribe Claderiinae Szlach. We publish the plastome and ITS sequences of Claderia viridiflora. Two names are lectotypified.
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Trias-Blasi, A.

Flora of Singapore precursors, 35: New records of Vitaceae in Singapore [Page 43 - 48]
Three species of Vitaceae are newly recorded for Singapore. One of them, Causonis maritima (Jackes) Jackes, is treated here as native, while two of them, Cissus discolor Blume and Cissus repanda Vahl, are presumed to be introduced.
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Turner, I. M.

Flora of Singapore precursors, 36: Notes on Pentaphragmataceae in Singapore [Page 49 - 53]
The nomenclature and typification of Pentaphragma Wall. ex G.Don (Pentaphragmataceae) and its Singapore representative are briefly reviewed. A lectotype is designated for the only generic synonym of Pentaphragma, Francfleurya A.Chev. & Gagnep. Four other lectotypifications for names of species or varieties are given, one at the second step.
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Turner, I. M.

Flora of Singapore precursors, 37: Notes on Goodeniaceae in Singapore [Page 55 - 59]
The nomenclature and typification of Scaevola taccada (Gaertn.) Roxb. and its synonyms are reviewed. In total, nine new lectotypifications are presented.
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Esser, H.-J.

Flora of Singapore precursors, 38: Additional new records of Euphorbiaceae for Singapore [Page 61 - 67]
Three species of Euphorbiaceae are newly recorded for Singapore, two of which (Euphorbia prostrata Aiton and Microstachys corniculata (Vahl) A.Juss. ex Griseb.) are non-native, and one (Croton erythrostachys Hook.f.) is presumed native but nationally extinct. The record of Microstachys corniculata is supposedly new for the continent. A lectotype is designated for Microstachys corniculata.
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Middleton, D.J.

Flora of Singapore precursors 39: Notes on Podocarpaceae in Singapore [Page 69 - 71]
The nomenclature and typification of the species of Podocarpaceae in Singapore are discussed.
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Saw, L.G.

A revision of the genus Iguanura (Arecaceae) in Peninsular Malaysia [ Page 73 - 117]
Eleven species of Iguanura are recognised for Peninsular Malaysia, one of them with two varieties. One species is described as new, Iguanura ruthiae Saw. All species are described, and notes on distribution, provisional IUCN conservation assessment and ecology are given. Thirteen taxa are synonymised and all names typified.
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Olimpos, S.M.B. & Ples, D.J.R.

Begonia dimorpha (Begoniaceae) and Clerodendrum pygmaeum (Lamiaceae), new records for the Philippines from Tawi-Tawi Island [Page 119 - 228]
Field work in Panglima Sugala, Tawi-Tawi, Philippines, resulted in new country records of Begonia dimorpha S.Julia (sect. Petermannia, Begoniaceae) and Clerodendrum pygmaeum Merr. (sect. Clerodendrum, Lamiaceae), both previously recorded only from Borneo. There are now 162 species of Begonia L. and 18 species of Clerodendrum L. known from the Philippines. Field observations and taxonomic notes on each species are presented and the presence of Bornean plant elements in the Sulu archipelago is discussed.
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Geiger, D.L.

Studies in Oberonia 11: The genus Hippeophyllum reduced to Oberonia, with ten new synonyms of Oberonia scortechinii (Orchidaceae: Malaxideae) [Page 129 - 148]
The genus Hippeophyllum Schltr. is synonymised under Oberonia Lindl. The following names are synonymised under Oberonia scortechinii Hook.f.: Hippeophyllum alboviride J.J.Sm., H. biakense J.J.Sm., H. celebicum Schltr., H. halmaherense J.J.Sm., O. hamadryas Ridl., O. longifolia Ridl., H. micranthum Schtr., H. papillosum Schltr., H. sulense J.J.Sm. and H. wenzelii Ames. Oberonia scortechinii is an epiphyte distributed from Malaysia through Indonesia, the Philippines, to New Guinea and the Solomon Islands at elevations of 0–500 m (rarely to 1000 m).
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Murugan, P., Ravichandran, V. & Murugan, C.

The genus Gynochthodes (Rubiaceae, Rubioideae, Morindeae) in India [Page 149 - 167]
The genus Gynochthodes Blume (Rubiaceae) in India is revised. Five species are recognised and described, including the new species Gynochthodes nilagiriensis P.Murugan, V.Ravich. & Murugan. Gynochthodes cochinchinensis (DC.) Razafim. & B.Bremer has been recorded for India but is excluded here. All names are typified.
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Karuppusamy, S., Ravichandran, V. & Bharath Simha Yadav, P.

Litsea megamalayana (Lauraceae), a new species from the southern Western Ghats of India [Page 169 - 175]
Litsea megamalayana Karupp., V.Ravich. & Bharath (Lauraceae), a new species from the Megamalai Wildlife Sanctuary of the southern Western Ghats, is described and illustrated. A detailed description, illustration, colour photographs and a table of comparisons to an allied species are provided here for easy identification and further study.
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Year of Publication 2022, Vol. 74 (2)

Date Published 28 December 2022
Turner, I.M. & Leong, P.K.F.

Pycnarrhena (Menispermaceae), a new generic record [Page 139 to Page 144]
A collection made from Nee Soon Swamp Forest in 2005 is confirmed as a specimen of Pycnarrhena fasciculata (Miers) Diels (Menispermaceae). This represents the first record of the genus Pycnarrhena for the native flora of Singapore.
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Ganesan, S.K.

Flora of Singapore precursors, 33: Further typifications and notes on Dipterocarpaceae [Page 145 to Page 149]
Nomenclatural notes on names of Dipterocarpaceae from Singapore are presented. Four names, including two synonyms, are lectotypified. These are Cotylelobium malayanum Slooten, Dipterocarpus apterus Foxw., Dipterocarpus kerrii King and Vatica maingayi Dyer. The etymology of Dipterocarpus kerrii is discussed.
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R.P.J. de Kok

Flora of Singapore precursors, 34: Typification of names in Singapore Acanthaceae [Page 151 to Page 158]
Nineteen names of Acanthaceae species occurring in Singapore are lectotypified and one name is neotypified
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Choo, L.M., Ang, W.F., Loo, A.H.B. & Er, K.B.H.

Unravelling the identity of Sindora (Fabaceae, Detarioideae) trees in thehistorical landscapes of Singapore [Page 159 to Page 181]
Sindora Miq. is a noteworthy genus in Singapore, with records of its existence, both natural and cultivated, dating back to the colonial period. Among which was the iconic ‘Changi Tree’, an approximately 76 m tall tree at Changi that was felled in 1942 during World War II. With the recent revision of the genus in Singapore, it was timely to survey the key historical landscapes at the Singapore Botanic Gardens (SBG), Fort Canning Park (FCP) and Changi Village Estate (CVE) for Sindora trees that are currently found or were once present in these locations. This was done through a site survey of the three areas, a study of herbarium specimens, literature and pictorial records relating to these areas, and the carbon dating of selected trees to ascertain their ages. At least 26 mature Sindora trees were recorded from this study, with at least 23 from SBG, one from FCP and two from CVE. Eighteen of these trees have been lost over time, leaving a total of eight trees consisting of an individual of Sindora siamensis Teijsm. ex Miq. and five of Sindora wallichii Benth. from SBG, one Sindora siamensis from FCP and one Sindora × changiensis L.M.Choo et al. from CVE. The latter is a hybrid of Sindora coriacea (Baker) Prain and S. echinocalyx Prain, recently elucidated using molecular tools. The identity of the ‘Changi Tree’ could not be fully verified, but it was likely to have been a Sindora echinocalyx based on a herbarium specimen collected from Changi. This study highlights the conservation value of trees in historical landscapes in Singapore, their importance in contributing to the genetic diversity of species in Singapore outside of nature reserves, and their role as a living legacy of Singapore’s rich botanical and horticultural history.
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Pannell, C.M.

Three new species in the Achariaceae from Peninsular Malaysia [Page 183 to Page 189]
One new species of Hydnocarpus Gaertn. and two new species of Ryparosa Blume from Peninsular Malaysia are described. Hydnocarpus perakensis Pannell is most similar to H. nanus King, but its fruits are larger and the leaves are hairy. Ryparosa suboppositifolia Pannell is most like R. acuminata Merr., but its leaves are not glaucous and it has an indumentum of
white medifixed hairs. Ryparosa reticulata Pannell is also most like R. acuminata Merr., but it differs in its indumentum of numerous adpressed orange-brown T-shaped hairs on the lower leaf surface, the leaves have up to 6 veins that are closer together and less steeply ascending,and the stamen filaments are free, whereas they form a tube in R. acuminata.
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Sulistiarini, D., Windadri, F.I., Sahroni, D., Sutikno & Surya, D.

The orchid diversity of Banggai Kepulauan, Central Sulawesi, Indonesia [Page 191 to Page 205]
Few orchid studies have focused on the satellite islands of Sulawesi, including the Banggai Kepulauan Archipelago. This study aims to understand the diversity of orchid species in Banggai Kepulauan and to add data on the diversity of orchids in Sulawesi. We identified 28 species of orchids from Banggai Kepulauan, of which two are endemic to Sulawesi: Dendrobium jubatum Schuit. & de Vogel and Dendrobium rhodobalion Schltr.; four species are only distributed in the eastern part of Indonesia: Cylindrolobus quadricolor (J.J.Sm.) Rauschert, Dendrobium lanceolatum Gaudich., Dendrobium purpureum Roxb. and Habenaria beccarii Schltr.; three species are new records for Sulawesi: Aphyllorchis acuminata J.J.Sm., Dendrobium acinaciforme Roxb. and Tainia trinervis (Blume) Rchb.f.; and one species is a new record for Banggai Kepulauan: Crepidium resupinatum (G.Forst.) Szlach.
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Rahayu, S., Ahmad R.P.P. & Rodda, M.

Hoya of Sulawesi, Indonesia: A checklist, two new species, a new subspecies and six new records [Page 207 to Page 221]
Two new Hoya R.Br. species and a new subspecies, Hoya towutiensis S.Rahayu, R.P.P.Ahmad & Rodda, Hoya mamasa S.Rahayu, R.P.P.Ahmad & Rodda and Hoya mamasa subsp. longicorolla S.Rahayu, R.P.P.Ahmad & Rodda, are described from Sulawesi, Indonesia. A checklist of Hoya of Sulawesi is published, including 30 species and one subspecies, six of
which are newly recorded. Hoya tsiangiana P.T.Li is typified and considered a new synonym of Hoya australis R.Br. ex J.Traill s.l. 
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Mustaqim, W.A., Hutabarat, P.W.K., Ahmad, R.P.P., Zulfadli & Ardi, W.H.

New and noteworthy records of spermatophytes in the Wallacea region [Page 223 to Page 242]
Some new and noteworthy records of spermatophytes in the Wallacea region are presented. These include three new species records in Wallacea for Lasianthus chrysotrichus Lauterb. (Rubiaceae), Medinilla medinilliana (Gaudich.) Fosberg & Sachet (Melastomataceae)and Thottea tomentosa (Blume) Ding Hou (Aristolochiaceae); new collection records for Pandorea pandorana (Andrews) Steenis (Bignoniaceae) for both Sulawesi and Buru (Maluku Archipelago); a new record in Sulawesi for Aeschynanthus amboinensis (Merr.) Mendum (Gesneriaceae); extended distributions in Sulawesi for six species: Ficus lawesii King (Moraceae), Gaultheria celebensis (J.J.Sm.) Kron & P.W.Fritsch (Ericaceae), Gaultheria hendrianiana (Argent) Kron & P.W.Fritsch (Ericaceae), Gaultheria retusa (Sleumer) Kron & P.W.Fritsch (Ericaceae), Calanthe stenocentron (Schltr.) M.W.Chase et al. (Orchidaceae) and Rhododendron torajaense Craven (Ericaceae); and an extended distribution in Maluku for Begonia aptera Blume (Begoniaceae). The morphology of the poorly documented Sulawesi endemic Gaultheria celebensis (J.J.Sm.) Kron & P.W.Fritsch is discussed.
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J. Leong-Škorničková, S. Soonthornkalump, S. Niwesrat & S.Q. Lim

Curcuma lindstromii (Zingiberaceae: Zingiberoideae), a new species from southeastern Thailand [Page 243 to Page 250]
Curcuma lindstromii Škorničk. & Soonthornk. (Zingiberaceae: Zingiberoideae), a new species from Thailand, is described. It is compared to the morphologically closest species from Curcuma subg. Ecomatae, Curcuma rhomba Mood & K.Larsen. A detailed description, a colour plate and information on its distribution, ecology, phenology and uses are provided. A provisional IUCN conservation assessment of Vulnerable is proposed for this species. A lectotype is designated for the name Curcuma stenochila Gagnep.
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Kurzweil, H., Tanaka, N., Mu Mu Aung & Ormerod, P.

A new species of Luisia (Orchidaceae) from Shan State, Myanmar [Page 251 to Page 256]
A new species, Luisia verrucosa Kurzweil, Nob.Tanaka & Ormerod, from Shan State in Myanmar is described and illustrated. It is very distinct in the genus, characterised by (1) light pinkish and purple-marked navicular sepals which are coarsely verrucose on the outside, especially in their distal part, (2) light pinkish and irregularly pink-dotted lorate-elliptic petals with yellow tips, and (3) a lip with a comparatively large, oblong-elliptic, maroon-purple epichile with prominent callosities in the form of massive bulges. Because its lip does not have a constriction or prominent boundary line between hypochile and epichile, it is superficially similar to Luisia brachystachys (Lindl.) Blume, but the new species differs by having considerably larger flowers with verrucose sepals and pronounced lip ornaments.
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Murugan, P. Kottaimuthu, R. & Murugan, C.

Typification of thirty-five names in Rubiaceae of the Southern Western Ghats, India [Page 257 to Page 274]
Found throughout the world, the Rubiaceae is the fourth largest family of angiosperms, with around 605 genera (POWO, 2021) and 13,000 species (Bremer, 2009). It is easily identified in the field by its simple, opposite or whorled, entire leaves, interpetiolar stipules, and gamopetalous flowers with an inferior ovary (Davis et al., 2009). In India, it is represented by 637 taxa (572 species, 14 subspecies, and 51 varieties) (Deb, 2001; Gangopadhyay et al., 2020) of which 130 species are endemic to the Southern Western Ghats (Nayar et al., 2014; Singh et al., 2015). During taxonomic studies on the Rubiaceae in South India, the authors found that 29 names had not yet been typified and a further six had only been partially typified. Therefore, these 35 names are here lectotypified, including six in a second step lectotypification.
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Jayasinghe, H.D., Wijesundara, D.S.A., Ranasinghe, R.A.S.W. & Kathriarachchi, H.S.

Two new species of Syzygium (Myrtaceae) from Sri Lanka, with lectotypification and recircumscription of Syzygium assimile
[Page 257 to Page 274]
Two new species of Syzygium Gaertn. from Sri Lanka, S. aureum Jayasinghe and S. hemachandrae Jayasinghe, are described. Syzygium aureum is distinguished from S. grande (Wight) Walp. in having a thick ring of nectarial glands near the rim of the hypanthium and in having subglobose fruits tinged with red at ripening. Syzygium hemachandrae differs from S. assimile Thwaites in having a longer petiole, an elliptic to ovate-elliptic lamina with fewer lateral veins, a cylindrical to subcylindrical peduncle at its basal parts and approximately quadrangular secondary axes. Syzygium aureum, which was previously misidentified as Syzygium firmum Thwaites (= S. grande), is restricted to the southwestern lowland rainforests, while S. hemachandrae is confined to the moist monsoon forests of the intermediate zone of eastern Sri Lanka. Both new species were found to be included amongst the syntypes of Syzygium assimile. We therefore designate a lectotype and recircumscribe Syzygium assimile.
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Year of Publication 2022, Vol. 74 (1)

Date Published 30 June 2022
Hughes, M.

Memecylon acuminatissimum, a new record for Singapore [Pg 1 - 3]
The tree species Memecylon acuminatissimum Blume is recorded as new to Singapore, with the only known locations being the Singapore Botanic Gardens’ Rain Forest and Chestnut Nature Park.
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Turner, I.M

Flora of Singapore precursors, 30: Notes on Symplocaceae in Singapore. [Pg 5 - 18]
The native Symplocaceae of Singapore (eight Symplocos species) are listed with synonymy and typification. A total of 30 lectotypifications are made, including six at the second step.
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Choo, L.M., Chen, L.M.J & Turner, I.M.

Flora of Singapore precursors, 31: The genus Boerhavia (Nyctaginaceae) in Singapore and clarification of Boerhavia diffusa in Singapore and Peninsular Malaysia [Pg 19 - 36]
Past studies of the genus Boerhavia in Singapore have listed Boerhavia diffusa L. as the only Boerhavia species with glandular anthocarps in Singapore. However, recent field collections and a study of herbarium specimens have uncovered the presence of two taxa hitherto included under the name ‘Boerhavia diffusa’ in Singapore. All material previously identified as Boerhavia diffusa in Singapore is Boerhavia repens L. More recently collected material, however, is the first actual material of Boerhavia diffusa for Singapore and is consequently a new record. A similar misapplication of the name Boerhavia diffusa also applies in Peninsular Malaysia. In light of this nomenclatural confusion pertaining to specimens in Singapore and Peninsular Malaysia, we clarify the use of the name Boerhavia diffusa L., resurrect the use of the name Boerhavia repens L., and provide a taxonomic revision and identification key for Boerhavia in Singapore.
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Ho, B.C. & Lua, H.K.

Flora of Singapore precursors, 32: Discoveries in Mucuna (Leguminosae, subfamily Papilionoideae) with a review of the genus in Singapore [Pg 37 - 55]
The discovery of Mucuna biplicata Teijsm. & Binn. ex Kurz in Singapore is an addition to the native flora. Mucuna pruriens (L.) DC. var. pruriens is reported as a non-native new record for Singapore. Mucuna gigantea (Willd.) DC. subsp. gigantea is rediscovered. With a total of four species, an identification key to the taxa of Mucuna in Singapore is provided. Descriptions are provided for all taxa based on the Singapore specimens. A lectotype is designated for Mucuna lucidula Burck.
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Middleton, D.J, Atkins, S., Beentje, H.J., Chen, L.M.J., Choo, L.M., de Kok R.P.J., de Wilde, W.J.J.O., Duyfjes, B.E.E., Ho, B.C., Lindsay, S. & Lua, H.K.

Additions to the Flora of Singapore: New and overlooked records of casual and naturalised plant species (6) [Pg 57 - 70]
Nine non-indigenous casual or naturalised species are newly recorded for Singapore: Justicia pectinata L. (Acanthaceae), Lepidagathis cephalotes (Link) Kuntze (Acanthaceae), Ruellia prostrata Poir. (Acanthaceae), Acmella ciliata (Kunth) Cass. (Asteraceae), Tilesia baccata (L.) Pruski (Asteraceae), Wollastonia asperrima Decne. (Asteraceae), Merremia gemella (Burm.f.) Hallier f. (Convolvulaceae), Momordica cochinchinensis (Lour.) Spreng. (Cucurbitaceae) and Verbena brasiliensis Vell. (Verbenaceae). Descriptions, notes on the distribution, and ecology are provided for all species, while colour
plates are provided for four species.
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R.P.J. de Kok

The Araliaceae of Peninsular Malaysia: five new species, two neotypifications and twenty-two lectotypifications in several genera [Pg 71 - 90]
One new species of Brassaiopsis, B. malayana de Kok, and four new species of Heptapleurum, H. frodinii de Kok, H. lengguanii de Kok, H. longicaudatum de Kok and H. nanocephalum de Kok, all Araliaceae from Peninsular Malaysia, are described here. Descriptions and notes on distribution, conservation status and ecology are given. In addition, 22 names in various genera of Araliaceae are lectotypified and two names are neotypified.
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Julia, S., Kiew, R. & Ling, C.Y.

Hexatheca longipedunculata (Gesneriaceae), a new species from Sarawak, Borneo [Pg 91 - 100]
Hexatheca longipedunculata S.Julia & Kiew, a new species from Sarawak, is described and illustrated. Hexatheca is a genus endemic to the island of Borneo. The description of this new species brings the number of species recognised in the genus to four. A key to the species and subspecies of Hexatheca and a distribution map of the newly described species are presented. Its conservation status is assessed.
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Rodda, M. & Rahayu, S.

Nine new species and one new subspecies of Hoya (Apocynaceae: Asclepiadoideae) from Borneo [Pg 101 - 129]
In the present paper we publish nine new species and one new subspecies from Borneo, Hoya ariffinii Rodda & S.Rahayu, H. boycei Rodda & S.Rahayu, H. curtisii King & Gamble subsp. collariata S.Rahayu & Rodda, H. dulitensis Rodda & S.Rahayu, H. kaikoeana S.Rahayu & Rodda, H. kapuasensis S.Rahayu & Rodda, H. kerangasensis Rodda & S.Rahayu, H. peltata Rodda & S.Rahayu, H. polypus S.Rahayu & Rodda and H. sangguensis S.Rahayu & Rodda. Five species are endemic to Kalimantan, two to Brunei and one to Sarawak. Only Hoya kerangasensis is found in Brunei, Sabah and Sarawak, and only H. sangguensis is also found outside Borneo in Peninsular Malaysia. With these new species the number of Hoya of Borneo reaches 85 species and four subspecies.
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Murugesan, M., Anusuba V., Arisdason, W., Tharani, R. & Karthik, B.

A new species of Arundinella (Poaceae) from the Nilgiri Biosphere Reserve, India [Pg 131 - 138]
Arundinella mukurthiana Murug. & Anusuba (Poaceae), a new species from the high-altitude grasslands of Mukurthi National Park, a part of Nilgiri Biosphere Reserve of the Western Ghats, Tamil Nadu, India, is described along with a detailed description, phenological information, a line drawing and photographs. A note on its conservation status is also provided.
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Year of Publication 2022, Vol. 74 (S1)

Date Published 21 May 2022

Dr David J. Middleton


 Dr Stuart Lindsay Dr Junhao Chen
     (Indexing)                              (Copy Editor)

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S. Lindsay, D.J. Middleton, B.C. Ho, K.Y. Chong, I.M. Turner, Ali Ibrahim, M. Alonso-García, W.F. Ang, P.S. Ashton, P. Athen, S. Atkins, Bazilah Ibrahim, H.J. Beentje, C.M. Boo, P.C. Boyce, G.L.C. Bramley, S. Buerki, M.W. Callmander, S. Chantanaorrapint, M. Cheek, C.-W. Chen, J. Chen, L.M.J. Chen, Chew, P.T., R. Chong, L.M. Choo, R.C.K. Chung, M.J.E. Coode, S.C. Chua, D. Cicuzza, R.P.J. de Kok, G.W.H. Davison, W.J.J.O. de Wilde, H. Duistermaat, A. Dubéarnès, B.E.E. Duyfjes, L.T. Ellis, H.-J. Esser, P.R. Gajurel, S.W. Gale, S.K. Ganesan, E.M. Gardner, D.L. Geiger, R.K. Harwood, Hassan Ibrahim, S. He, A. Henderson, P.H. Hovenkamp, M. Hughes, Zaki Jamil, M.H.P. Jebb, D.M. Johnson, A. Kartonegoro, R. Kiew, S. Knapp, S.L. Koh, H. Kurzweil, S. Lee, P.K.F. Leong, J. Leong-Škorničková, G.A. Levin, D.C.H. Liew, R.C.J. Lim, W.H. Lim, A.H.B. Loo, Y.W. Low, H.K. Lua, S. Lum, D.J. Mabberley, R. Mahyuni, B. Maslin, N.A. Murray, L. Neo, X.Y. Ng, K.M. Ngo, M.A. Niissalo, P.T. Ong, C.M. Pannell, A. Phang, G.T. Prance, C. Promma, C. Puglisi, M.L. Rodda, S.C.K. Rubasinghe, R.M.K. Saunders, I.A. Savinov, L.G. Saw, A. Schuiteman, W.W. Seah, D.A. Simpson, J.S. Strijk, P. Sukkharak, M. Sugumaran, S. Syahida- Emiza, J.P.C. Tan, N.P. Taylor, Y.K.L. Teo, D.C. Thomas, A. Trias-Blasi, T. Utteridge, P.C. van Welzen, J.F. Veldkamp, J. Vermeulen, R. Wang, P. Wilkie, Y.-M. Wei, S.Y. Wong, K.M. Wong, S. Yaakub, T.W. Yam, S. Yang, T.L. Yao, W. Ye, A.T.K. Yee, C.K. Yeo, Y.S. Yeoh, C. Yong, K.T. Yong, N.J.C. Zerega, R.-L. Zhu, K.B.H. Er

Flora of Singapore: Checklist and bibliography [Page 3 - 860]
A checklist of all species of bryophytes, lycophytes, ferns, gymnosperms and angiosperms that are found in the wild (native, naturalised and casual) in Singapore is presented. We have attempted to account for all names of species and infraspecific taxa that have ever been recorded for Singapore, along with the pertinent publications that reported each of these names. For each currently accepted name, the synonyms of relevance for Singapore are included. The native or non-native status for all taxa is given, along with the most recent national conservation assessment applied to each native taxon. If we were aware that the most recent assessment required an update, the taxon is newly assessed here. The checklist includes 2654 native taxa, 479 naturalised/casual taxa and 101 cryptogenic taxa.
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