Year of Publication: 2016, Vol. 68 (2)

Date Published 15 November 2016
M. Rodda, N. Simonsson Juhonewe & D.J. Middleton
The taxonomic status of the presumed extinct Singaporean Hoya wallichii (Apocynaceae: Asclepiadoideae) [Page 175 - 187]
After critical study of type material and all available collections, the identity of Hoya wallichii (Wight) C.M.Burton as a species distinct from Hoya campanulata Blume is clarified. Hoya wallichii was previously considered to be endemic to Singapore but had become nationally extinct and consequently globally extinct. This study reveals, however, that it is also found in Peninsular Malaysia and Brunei and, even though it is still nationally extinct in Singapore, it is no longer globally extinct. Hoya campanulata is widespread and locally common in Sundaland but also nationally extinct in Singapore. The two species are fully described and illustrated by line drawings and colour photographs. Two conservation assessments are made, three names are lectotypified, and one name is epitypifed. This paper exemplifies how critical taxonomic understanding is fundamental to meaningful conservation assessments.

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R.P.J. de Kok, S. Sengun & G.L.C. Bramley
Two new records for the Lamiaceae of Singapore [Page 189 - 200]
The species of Lamiaceae have recently been revised for Singapore. In total 44 species in 21 genera are recorded, of which 23 species are native, an additional five are probably native, and 16 are non-native species which have naturalised in Singapore. One new record for Singapore has been found (Vitex rotundifolia) and one species reinstated (Callicarpa pentandra) after being ignored for over a hundred years.

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R.P.J. de Kok
Notes on the monotypic genus Hexapora (Lauraceae), endemic to Peninsular Malaysia [Page 201 - 208]
An account of the genus Hexapora (Lauraceae) is presented. It comprises morphological descriptions of the genus and species, a provisional IUCN conservation assessment, ecological information and taxonomic notes. The morphology of the genus is compared to that of a number of possible close relatives, leading to the conclusion that it most likely belongs within the Beilschmiedia clade, and is likely closely related to Sinopora.
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J. Leong-Škorničková & R. Kiew
Hanguana fraseriana (Hanguanaceae), a new species from Peninsular Malaysia [Page 209 - 214]
Hanguana fraseriana, a new species from Peninsular Malaysia, is described and illustrated here.

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T.L. Yao
Nomenclatural updates and new records in Peninsular Malaysian Pentaphylacaceae [Page 215 - 223]
Four nomenclatural updates, viz. Ternstroemia coriacea, T. palembangensis, T. penangiana and T. wallichiana, and two new records T. patens and Eurya glabra in Peninsular Malaysia are presented.

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R. Kiew
Impatiens (Balsaminaceae) species from karst limestone in Kelantan, Malaysia, including three new species [Page 225 - 238]
Peninsular Malaysian Impatiens foxworthyi M.R.Hend. is distinct from the Thai I. opinata Craib. Three new Impatiens species are described: I. glaricola Kiew with purple flowers, I. chikuensis Kiew with pale yellow flowers, and I. vinosa Kiew with deep red flowers. While Impatiens foxworthyi is widespread on karst limestone in Kelantan and Pahang, the three new species are narrowly endemic to Kelantan limestone and are critically endangered.

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R. Kiew, S. Julia & C.Y. Ling
The Calcarea group of Begonia (Begoniaceae) from Borneo, including four new species from Sarawak [Page 239 -255]
Four new species, Begonia kanaensis Kiew & C.Y.Ling, Begonia lingiae S.Julia, Begonia rubrotepala S.Julia and Begonia sadirensis Kiew & S.Julia are described from Sarawak, Borneo. Notes on habitat, distribution and additional specimens examined for Begonia calcarea and B. sabahensis are also provided. A key to the species in the ‘calcarea group’ is provided. The status of the ‘calcarea group’ as belonging to Begonia sect. Diploclinium or to a new section is discussed. A provisional conservation status is assigned to each species.

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S. Julia & R. Kiew
Eight new Begonia (Begoniaceae) species from the Lanjak Entimau Wildlife Sanctuary and Batang Ai National Park, Sarawak, Borneo [Page 257- 277]
Eight new Begonia (Begoniaceae) species, Begonia addrinii S.Julia & Kiew, Begonia celata S.Julia & Kiew, Begonia crassa S.Julia & Kiew, Begonia devexa S.Julia & Kiew, Begonia fractiflexa S.Julia & Kiew, Begonia hirtitepala S.Julia & Kiew, Begonia jamilahanuiana S.Julia and Begonia ubahribuensis S.Julia & Kiew, are described from Lanjak Entimau Wildlife Sanctuary and Batang Ai National Park in Sarawak, Malaysia. Two and four species are only known from Batang Ai National Park and Lanjak Entimau Wildlife Sanctuary respectively while two species are found in both sites. Seven species belong to Begonia sect. Petermannia and the placement of one (Begonia fractiflexa) is uncertain.

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N.K.E. Undaharta & W.H. Ardi
Studies on Begonia (Begoniaceae) of the Moluccas III: A new Begonia from Seram, Indonesia [Page 279 - 285]
A new species of Begonia, Begonia nephrophylla Undaharta & Ardi, is described from Manusela National Park, Seram Island, Moluccas, Indonesia. The species is endemic to Seram and belongs to Begonia section Petermannia. An illustration of the new species and a key to the Moluccan species of Begonia is presented.

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J. Leong-Škorničková, Q.B. Nguyễn, H.Đ. Trần & E. Záveská
Etlingera poulsenii and Hornstedtia bella (Zingiberaceae: Alpinieae), two new species from central Vietnam [Page 287 - 297]
Two new ginger species from central Vietnam, Etlingera poulsenii Škorničk. and Hornstedtia bella Škorničk., are described and illustrated here.

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J.D. Mood, N. Tanaka, M.M. Aung & J. Murata
The genus Boesenbergia (Zingiberaceae) in Myanmar with two new records [Page 299 - 318]
The taxonomic history of Boesenbergia Kuntze (Zingiberaceae) in Myanmar is reviewed. Based on specimen records eight species are currently confirmed as occurring in Myanmar. These include two new records, Boesenbergia albomaculata S.Q.Tong and B. kerrii Mood, L.M.Prince & Triboun. Two previously listed species, Boesenbergia plicata (Ridl.) Holttum and B. thorelii (Gagnep.) Loes., are not considered here due to lack of specimens originating in Myanmar. A key to the species is provided with a description of each based on living material.

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K. Karthigeyan, M. Das Das & L.S. Wijedasa
Memecylon cerasiforme (Melastomataceae): a poorly known species rediscovered, redescribed and newly recorded for India [Page 319 - 326]
The species Memecylon cerasiforme Kurz was previously known only from the type which was collected in Chittagong, Bangladesh. A recent collection of the species from the Jaldapara National Park, West Bengal, India is not only the first since the species was described in 1877 but also the first record of its occurrence in India and the first flowering collection. Its flowers can now be described and illustrated for the first time. An emended description, figure, phenological and habitat information, notes on related species, and a conservation assessment are also provided. The only original material available at Kew is designated here as a lectotype.

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David J. Middleton
Book Review [Page 327 - 328]
Flora Malesiana, series I – Seed Plants. Volume 22: Lythraceae. W.J.J.O. de Wilde & B.E.E. Duyfjes. P.C. van Welzen (Ed.) 2016.

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Year of Publication: 2016, Vol. 68 (1)

Date Published 20 May 2016
D.J. Middleton
A revision of Aeschynanthus (Gesneriaceae) in Singapore and Peninsular Malaysia [Page 1 - 63]
The genus Aeschynanthus Jack is revised for Singapore and Peninsular Malaysia. Four species for Singapore and fourteen species for Peninsular Malaysia are recognised, keys to the species are given, all names are typified, and detailed descriptions of all species are provided. Conservation assessments are provided for all species. Eleven names are lectotypified here and one epitype is designated.

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I.M. Turner
Notes on the Annonaceae of the Malay Peninsula [Page 65 - 69]
Notes concerning the three genera Alphonsea Hook.f. & Thomson, Artabotrys R.Br. ex Kew Gawl. and Stelechocarpus Hook.f. & Thomson (Annonaceae) for the Malay Peninsula are presented. Alphonsea kingii J.Sinclair is accepted as an endemic species and a second-stage lectotypification is presented for Alphonsea cylindrica King. Seven species of Artabotrys are lectotypified. The recently described genus Winitia Chaowasku is reduced to Stelechocarpus and a new combination is made for Winitia expansa Chaowasku.

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K.M. Wong, A.K. Muhammad Ariffin & A.A. Joffre
Novitates Bruneienses, 5. Polyalthia watui (Annonaceae), a new tree species from Brunei, Borneo [Page 71 - 76]
A new species, Polyalthia watui, is described. It resembles Polyalthia bullata King in having bullate leaves with a cordate base, but is distinguished by its flowers with shorter pedicels, longer sepals and petals, subsessile monocarps with long stiff erect hairs; and broad-obovate leaves with fewer pairs of secondary veins. The new species is only known from Brunei’s Belait and Tutong districts and is very likely endemic to the Belait geosyncline that includes these areas.

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D. Girmansyah
Three new species of Begonia (Begoniaceae) from Sumbawa Island, Indonesia [Page 77 - 86]
Three new species of Begonia L., Begonia sumbawaensis Girm., Begonia brangbosangensis Girm. and Begonia jaranpusangensis Girm., are described from Sumbawa, Indonesia. All species are illustrated and identification keys and distribution maps are provided.

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D. Sulistiarini, D. Arifiani & Y. Santika
New records of Orchidaceae from Bali, Indonesia [Page 87 - 95]
During plant inventory work in the forested areas of Mt Mesehe and Mt Merbuk, Jembrana district, Bali, twenty orchid species were collected. Among the species collected, five were confirmed as new records for Bali. The five newly recorded species are Calanthe zollingeri Rchb.f., Plocoglottis plicata (Roxb.) Ormerod, Podochilus serpyllifolius (Blume) Lindl., Styloglossum speciosum (Blume) T.Yukawa & P.J.Cribb, and Vanilla albida Blume. Descriptions of taxa and images are provided.

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R. Kiew
Olea luzonica (Oleaceae), a new name for O. obovata from the Philippines [Page 97 - 98]
A new name, Olea luzonica Kiew, is proposed to replace the later homonym Olea obovata (Merr.) Kiew.

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T. Phutthai & M. Hughes
A new species and a new record in Begonia sect. Platycentrum (Begoniaceae) from Thailand [Page 99 -107]
Begonia khaophanomensis Phutthai & M.Hughes is described from a collection made on granitic rock in Krabi Province in Peninsular Thailand. It is endemic to the typelocality of Khao Phanom Bencha National Park, and is assessed as Least Concern using the IUCN criteria. Begonia macrotoma Irmsch. is noted as a new record for Thailand from collections in Chiangmai, Nakhon Ratchasima and Nakhon Nayok provinces; the species is also found in China, Myanmar, NE India, Nepal and Vietnam. Both species belong to Begonia sect. Platycentrum. A key to the species of Begonia sect. Platycentrum in Thailand is provided.

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J.D. Mood, A.G. Hussain & J.F. Veldkamp
The resurrection of Boesenbergia albosanguinea (Zingiberaceae) with a new record for Peninsular Thailand [Page 109- 124]
Boesenbergia albosanguinea (Ridl.) Loes. (Zingiberaceae) is resurrected from the synonymy of B. prainiana (King ex Baker) Schltr. It is redescribed in detail with comparative tables, figures, and illustrations. This species is a new record for Thailand.

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J.D. Mood, H.Đ. Trần, J.F. Veldkamp & L.M. Prince
Boesenbergia siphonantha (Zingiberaceae), a new record for Thailand and Vietnam with notes on the molecular phylogeny [Page 125 - 137]
Boesenbergia siphonantha (King ex Baker) M.Sabu, Prasanthk. & Škorničk. (Zingiberaceae) is newly recorded from Thailand and Vietnam. Its nomenclatural history, taxonomy, and molecular phylogeny are discussed. Figures, a comparative table and a watercolour illustration are provided.

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F. Wen, T.V. Do, X. Hong, S. Maciejewski & Y.G. Wei
Boeica ornithocephalantha (Gesneriaceae), a new species from northern Vietnam [Page 139 - 144]
A new species, Boeica ornithocephalantha F.Wen, Y.G.Wei & T.V.Do (Gesneriaceae), is described from the northern part of Vietnam. This new species is most similar to Boeica ferruginea Drake, but clearly differs from the latter by the unusual corolla. A provisional IUCN conservation assessment is provided.

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M. Möller, K. Nishii, H.J. Atkins, H.H. Kong, M. Kang, Y.G. Wei, F. Wen, X. Hong & D.J. Middleton
An expansion of the genus Deinostigma (Gesneriaceae) [Page 145 - 172]
Based on molecular, morphological and cytological studies the previously monotypic genus Deinostigma W.T.Wang & Z.Y.Li has been expanded to include several species previously ascribed to Primulina Hance. Deinostigma now comprises seven species, including one previously placed in synonymy. The new combinations Deinostigma cicatricosa (W.T.Wang) D.J.Middleton & Mich.Möller, Deinostigma cycnostyla (B.L.Burtt) D.J.Middleton & H.J.Atkins, Deinostigma cyrtocarpa (D.Fang & L.Zeng) Mich.Möller & H.J.Atkins, Deinostigma eberhardtii (Pellegr.) D.J.Middleton & H.J.Atkins, Deinostigma minutihamata (D.Wood) D.J.Middleton & H.J.Atkins and Deinostigma tamiana (B.L.Burtt) D.J.Middleton & H.J.Atkins are made. Deinostigma eberhardtii is lectotypified. The genus is defined by a combination of an alternate leaf arrangement, hooked hairs on many plant parts, flowers with the pedicel inserted at an angle and off-centre on the receptacle, and, where known, a somatic chromosome number (2n) of < 36. This new circumscription of the genus expands its distribution from Vietnam into South China.
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J.F. Veldkamp
The valid publication of Monolophus (Zingiberaceae) revisited [Page 173 - 174]
The nomenclatural history of Monolophus Delafosse, Guill. & J.Kuhn is discussed. The combinations Monolophus picheansoonthonii (Phokham & Prasarn) Veldk. and Monolophus pubescens (Picheans. & Phokham) Veldk. are made.
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Year of Publication: 2015, Vol. 67 (2)

Date Published 23 December 2015
R.P.J. de Kok
Cryptocarya nitens (Lauraceae), a new species record for Singapore [Page 253 - 259]
Cryptocarya nitens (Blume) Koord. & Valeton is newly recorded for Singapore.  It was discovered during surveys in Bukit Timah Nature Reserve and the Nee Soon Swamp Forest. A description is given, together with a key based mostly on vegetative characters for all Cryptocarya species occurring in Singapore. A short overview of the Lauraceae of Singapore shows that, in total, 57 species belonging to 14 genera have been recorded, of which 47 species in 13 genera are native. Cryptocarya nitens is lectotypified in addition to two of its synonyms.

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Y.Y. Sam
Boesenbergia basispicata (Zingiberaceae), a new record for Peninsular Malaysia [Page 261 - 265]
Boesenbergia basispicata K.Larsen ex Sirirugsa was recently discovered from a limestone hill in Perak, Peninsular Malaysia. This brings the total number of Boesenbergia species in Peninsular Malaysia to 12. A description, colour plates and notes on this species are provided.

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K.M. Wong
Schizostachyum kuisingii, a new species of bamboo (Poaceae: Bambusoideae) from Peninsular Malaysia [Page 267 - 274]
Schizostachyum kuisingii K.M.Wong is a new species of bamboo from Peninsular Malaysia, distinguished from closely related species from adjacent areas in Indonesia, such as Schizostachyum castaneum Widjaja, Schizostachyum lutescens Widjaja and Schizostachyum mampouw Widjaja, by the hairs of the culm internode, glabrescent foliage leaves, leaf ligule form, a palea longer than the lemma and the presence of two lodicules in the flower.

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K.M. Wong, Y.W. Low, A.K. Muhammad Ariffin & A.A. Joffre
Novitates Bruneienses, 4. New records in the Araliaceae, Araucariaceae, Arecaceae, Fagaceae, Musaceae and Thymelaeaceae [Page 275 - 287]
Aralia merrillii C.B.Shang (Araliaceae), Agathis lenticula de Laub. (Araucariaceae), Licuala collina Saw and L. miriensis Saw (Arecaceae), Lithocarpus bullatus Hatus. ex Soepadmo and L. hallieri (Seemen) A.Camus (Fagaceae), Musa lawitiensis var. suratii (Argent) Häkkinen (Musaceae) and Aquilaria microcarpa Baill. (Thymelaeaceae) are newly recorded for the Brunei flora.

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T.L. Yao
Three new species of Loxocarpus (Gesneriaceae) from Sarawak, Borneo [Page 289 - 296]
Three new species, Loxocarpus burttii T.L.Yao, Loxocarpus littoralis T.L.Yao and Loxocarpus segelamensis T.L.Yao are described from Sarawak, Malaysia.
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W.H. Ardi & D.C. Thomas
Studies on Begonia (Begoniaceae) of the Moluccas II: A new species from Seram, Indonesia [Page 297 - 303]
A new species of Begonia L., Begonia galeolepis Ardi & D.C.Thomas, is described from Seram, Maluku province, Indonesia. The species is endemic to Seram and belongs to Begonia section Petermannia. An identification key to the seven Begonia species known from the Moluccas is provided.

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M. Rodda
A new typification of Hoya zollingeriana (Apocynaceae, Asclepiadoideae) [Page 305 - 307]
The typification of Hoya zollingeriana Miq. is discussed and a lectotype is selected from the original material. An earlier typification was based on a specimen belonging to a different taxon and is rejected.

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R.P.J. de Kok
A revision of Cryptocarya (Lauraceae) from Thailand and Indochina [Page 309 - 350]
A revision of the species of Cryptocarya R.Br. (Lauraceae) from Thailand and Indochina (Laos, Cambodia and Vietnam) is presented, comprising a key to species, full descriptions, distribution maps, provisional IUCN conservation assessments, ecological information and ethno-botanical notes where appropriate. In this treatment, 16 species are recognised, one species name is validated (Cryptocarya globularia Kosterm. ex de Kok), nineteen names are lectotypified and nine names are placed into synonymy for the first time. Using standard IUCN criteria, most species are considered to be Least Concern, one is assessed as Data Deficient (C. globularia), and six are Endangered (Cryptocarya chanthaburiensis Kosterm., Cryptocarya hainanensis Merr., Cryptocarya laotica (Gagnep.) Kosterm., Cryptocarya pallens Kosterm., Cryptocarya pustulata Kosterm. and Cryptocarya sublanuginosa Kosterm.).

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H.T. Lưu, J. Leong-Škorničková, L.X.B. Nguyễn, C.T. Đỗ & T.T. Hoàng
Newmania sessilanthera (Zingiberaceae): a new species from Vietnam [Page 351 - 355]
Newmania sessilanthera Lưu & Škorničk., a new species from Phú Yên province in central Vietnam, is described and illustrated here. Notes on this species, the third described to date, and a new key to the species of Newmania N.S.Lý & Škorničk. are provided.
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P.S. Ashton
A few last words on Far Eastern Dipterocarps [Page 357 -360]
Three additions to the Dipterocarpaceae are presented. Recent collections from southern Vietnam have finally revealed the identity of Dipterocarpus condorensis Pierre, necessitating reduction of D. caudatus Foxw. and nomenclatural adjustment of its subspecies; an unexpected novelty, Dipterocarpus pseudocornutus P.S.Ashton, is described from Tawi-Tawi, SW Philippines; while a new lower montane point endemic, Vatica palongensis P.S. Ashton, is reported from West Kalimantan.

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D.M. Johnson & N.A. Murray
A contribution to the systematics of Xylopia (Annonaceae) in Southeast Asia [Page 361 - 386]
Herbarium and field study of Xylopia L. (Annonaceae) for the Flora of Peninsular Malaysia and the Flora of Thailand projects has clarified regional diversity patterns within this ecologically significant lowland rainforest genus. Two species groups represented within Southeast Asian floras are delineated, one centred on Xylopia ferruginea (Hook.f. & Thomson) Baill. and the other on Xylopia malayana Hook.f. & Thomson. In the Xylopia ferruginea group, a new species, Xylopia erythrodactyla D.M.Johnson & N.A.Murray, is distinguished from X. ferruginea, and a new combination, Xylopia sumatrana (Miq.) D.M.Johnson & N.A.Murray, is proposed, based on an earlier name for the species currently known as Xylopia stenopetala Oliv. In the Xylopia malayana group, review of the species Xylopia elliptica Maingay ex Hook.f. & Thomson resulted in the recognition of three additional species: Xylopia platycarpa D.M.Johnson & N.A.Murray, from southern Thailand and northwestern Peninsular Malaysia, Xylopia ngii D.M.Johnson & N.A.Murray, from Peninsular Malaysia, Sumatra and Borneo, and Xylopia heterotricha D.M.Johnson & N.A.Murray, from Sumatra and Borneo. The taxon Xylopia malayana Hook.f. & Thomson var. obscura Kochummen is placed in synonymy under Xylopia elliptica sensu stricto. Xylopia fusca Maingay ex Hook.f. & Thomson var. sessiliflora Kochummen & Whitmore is distinguished from Xylopia fusca, and raised to species status as Xylopia sessiliflora (Kochummen & Whitmore) D.M.Johnson & N.A.Murray. We recognise 23 Xylopia species in the Sundaic region of Southeast Asia, and provide evidence that additional collecting and taxonomic analysis in the region is needed.

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K.M. Wong
Studies in Southeast Asian Melastoma (Melastomataceae), 1. Morphological variation in Melastoma malabathricum and notes on rheophytic taxa and interspecific hybridisation in the genus [Page 387 - 401]
Morphological variation in the widespread Indo-Pacific Melastoma malabathricum L. is discussed, as well as the possibility of hybridisation or introgression with other species. Hypanthium indumentum is a reliable character to diagnose the group and species. Five rheophytic taxa, including four distinctive species and a variant of the widespread M. malabathricum, which are superficially alike because of their stenophyllous leaf form, are enumerated for Southeast Asia and a key is provided for their identification. Melastoma kahayanense K.M.Wong from Borneo is described for the first time and the Sumatran Melastoma stenophyllum Merr. is considered a distinct species and not synonymous to M. malabathricum. Various putative hybrids or introgression products involving M. malabathricum are identified.

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I.M. Turner
Clarifying the nomenclature of Crateva trifoliata (Capparaceae) [Page 403 - 406]
Capparis trifoliata Roxb., the supposed basionym of ‘Crateva adansonii subsp. trifoliata (Roxb.) Jacobs’ and ‘Crateva trifoliata (Roxb.) B.S.Sun’, is shown to represent an illegitimate renaming of Crateva religiosa G.Forst. However, Jacobs effectively excluded the nomenclatural type of Capparis trifoliata, thus avoiding creating a superfluous name, and instead described Crateva adansonii subsp. trifoliata Jacobs validated by the citation of the Latin description of Crateva erythrocarpa Gagnep. Sun also excluded the type of Crateva religiosa, but his effective transfer of C. adansonii subsp. trifoliata to specific rank does not have priority over Crateva erythrocarpa Gagnep., which is therefore the correct name for this taxon when it is considered a distinct species. The name Crateva nurvula Buch.-Ham. is formally lectotypified.

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D.C. Thomas
BOOK REVIEW [Page 407 - 408]
Book Review:
A Guide to Begonias of Borneo. Ruth Kiew, Julia Sang, Rimi Repin & Joffre Ali Ahmad. 2015.

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D.J. Middleton 
BOOK REVIEW [Page 409 - 410]
Book Review:
Forest Trees of Southern Thailand, Volume 1 (A–Es). Simon Gardner, Pindar Sidisunthorn & Kongkanda Chayamarit. 2015.

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Year of Publication: 2015, Vol. 67 (01)

Date Published 04 June 2015
J. Leong-Škorničková & P.C. Boyce
Hanguana in Singapore demystified: an overview with descriptions of three new species and a new record [Page 1 - 28]
The genus Hanguana (Hanguanaceae – Commelinales) was until recently believed to be represented in Singapore by a single species, Hanguana malayana. Recent extensive surveys, coupled with the detailed study of fresh and herbarium material, however, suggests the presence of six species. In addition to the recently described Hanguana neglecta, three additional species, H. corneri, H. rubinea and H. triangulata, are described here as new to science. Of the three newly described species, Hanguana rubinea and H. triangulata are native, while H. corneri is believed to have been introduced from Peninsular Malaysia to Singapore and planted in Bukit Timah Nature Reserve in the 1930s. Hanguana nitens is recorded for the first time from Singapore and is also certainly native. While a large helophytic species of Hanguana, currently interpreted to be Hanguana malayana, is cultivated as an ornamental plant in Singapore, no native populations have been observed by us in the wild and neither do any herbarium records exist to suggest that this species was ever native to Singapore. Colour plates and a key to all Hanguana species are provided. Notes on habitats and preliminary IUCN assessments are also included for all native species to better facilitate conservation efforts of Hanguanaceae in Singapore. The existence and taxonomic potential of scale structures sheathing the inner staminodes in female flowers, named here as staminodial scales, are highlighted here for the first time. The importance of seed characters is also discussed.
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M. Rodda, D. Lee & S. Mishak
Dischidia acutifolia (Apocynaceae, Asclepiadoideae) a new record for the Singapore flora [Page 29 - 33]
We report the discovery of Dischidia acutifolia Maingay ex Hook.f., a new record for the Singapore flora. The species is only known from a single locality in Sungei Buloh Wetland Reserve and is therefore considered Critically Endangered in Singapore. A lectotype for the name is selected.
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I.M. Turner
From Ancistrocladus to Tristaniopsis via Tetramerista - the convoluted history of a Wallich collection and its impact on the native flora of Singapore [Page 35 - 37]
Ancistrocladus sagittatus Wall. ex Planch. (Ancistrocladaceae) is the name of a plant species solely based on a sterile collection made by Nathaniel Wallich in Singapore in 1822. Since 1875, this name has been treated as a variety or synonym of Tetramerista glabra Miq. (Tetrameristaceae). However, the original material is actually referable to Tristaniopsis (Myrtaceae), and Ancistrocladus sagittatus is here lectotypified and reported as a synonym of Tristaniopsis merguensis (Griff.) Peter G.Wilson & J.T.Waterh. for the first time. The new determination of the Wallich collection means that the record of Tetramerista glabra in the native flora of Singapore requires reassessment. It is concluded that Tetramerista glabra, the genus Tetramerista and the family Tetrameristaceae should all be excluded from inventories of the native flora of Singapore.
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S. Lindsay & D.J. Middleton
The clarification and synonymisation of two taxa of Vittaria from Peninsular Malaysia and a new combination in Haplopteris (Pteridaceae subfam. Vittarioideae) [Page 39 - 43]
A variety and a species of Vittaria in Peninsular Malaysia are synonymised and the new combination Haplopteris sessilifrons (Miyamoto & H.Ohba) S.Linds. is made.
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A.R. Rafidah, S. Syahida-Emiza, R. Kiew & K. Imin
Bothriospermum (Boraginaceae), a new generic record for the Flora of Peninsular Malaysia [Page 45 - 49]
Bothriospermum zeylanicum (J.Jacq.) Druce (Boraginaceae), a species native to Asia, has recently become naturalised in Cameron Highlands, Pahang, Malaysia. A description of the genus and species together with colour photographs are provided. It is the only representative of Boraginaceae sensu stricto (i.e. subfam. Boraginoideae) in Peninsular Malaysia.
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A.A. Joffre, A.K. Muhammad Ariffin, Y.W. Low & K.M. Wong
Novitates Bruneienses, 1. A background to the botanicalsurvey of Brunei Darussalam, and a new species of Jarandersonia (Malvaceae) [Page 51 - 60]
A brief introduction to research and botanical documentation of the Brunei flora, and the collaborative programme for a continuing botanical survey of the country, is given. An outline of the key biogeographical features of the Brunei area supports the premise that distinct geo-ecological enclaves occur which are special units of species richness, within which a significant level of undocumented plant diversity still exists. Jarandersonia yahyantha K.M.Wong, Joffre, Ariffin & Y.W.Low (Malvaceae), a new tree species so far only known in Brunei, is described.
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Y.W. Low, A.A. Joffre & A.K. Muhammad Ariffin
Novitates Bruneienses, 2. A remarkable new species of Begonia sect. Petermannia (Begoniaceae) from Brunei Darussalam [Page 61 - 68]
Begonia jamilahana Y.W.Low, Joffre & Ariffin is described and illustrated here as a new species based on a collection from Ladan Hills Forest Reserve, Tutong, Brunei Darussalam. This new taxon is closely related to B. conniegeriae S.Julia & Kiew and B. papyraptera Sands, but differs in a suite of morphological characters. Begonia jamilahana is placed in Begonia section Petermannia, the largest of all the sections recognised.
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K.M. Wong & Y.W. Low
Novitates Bruneienses, 3. Eight new woody plants in the Brunei flora, including five new species [Page 69 - 84]
Five new species of seed plants are described: Alangium kayuniga K.M.Wong (Cornaceae), Melastoma ariffinii K.M.Wong, Melastoma ashtonii K.M.Wong, Melastoma joffrei K.M.Wong & Y.W.Low (Melastomataceae), and Lasianthus jangarunii Y.W.Low (Rubiaceae). New plant records for the ongoing Brunei floristic inventory include Planchonea valida (Blume) Blume (Lecythidaceae), Melastoma velutinosum Ridl. (Melastomataceae), and Gardenia costulata Ridl. (Rubiaceae).
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S. Rahayu, U. Meve & M. Rodda
Hoya undulata (Apocynaceae, Asclepiadoideae), a new myrmecophytic species from Borneo, and typification of H. darwinii [Page 85 - 94]
In this paper we document the discovery of a new Hoya species from Borneo, H. undulata S.Rahayu & Rodda. The leaves of the new species form domatia, specialised structures harbouring ants, previously only observed in Hoya mitrata Kerr and H. darwinii Loher. The corona is unique among Bornean Hoya species as its lobes develop two lateral hooked appendages, also present in H. griffithii Hook.f, a species from mainland Asia. A lectotype for Hoya darwinii is also selected.
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N.K.E. Undaharta, I Made Ardaka, Agung Kurniawan & Bayu Adjie
Begonia bimaensis, a new species of Begonia from Sumbawa Island, Indonesia [Page 95 - 99]
A new species of Begonia sect. Reichenheimea, B. bimaensis Undaharta & Ardaka, is described from Mt Muria in Bima District, Sumbawa Island, Indonesia. The species is a narrow endemic, currently known from only one population, and has been assessed as Vulnerable using IUCN criteria.
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C. Puglisi, S. Suddee, P. Triboun & D.J. Middleton
A new species of Paraboea (Gesneriaceae) from Thailand [Page 101 - 106]
A new species, Paraboea maculata C.Puglisi, is described. It is compared to similar species and the habitat on granite rocks, unusual for the genus, is highlighted.
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H. Kurzweil & Saw Lwin
New orchid records for Myanmar, including the first record of the genus Stereosandra [Page 107 - 122]
Nine orchid species, Bulbophyllum capnophyton J.J.Verm. et al., Dendrobium macrostachyum Lindl., D. praecinctum Rchb.f., Habenaria reniformis (D.Don) Hook.f., Liparis distans C.B.Clarke, L. resupinata Ridl., Micropera thailandica (Seidenf. & Smitinand) Garay, Oberonia obcordata Lindl. and Stereosandra javanica Blume, are reported as new for Myanmar. Of particular interest is the Stereosandra as it is the first record of this genus in the country.
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M.F. Newman
A new species of Zingiber (Zingiberaceae) from Lao P.D.R. [Page 123 - 127]
Zingiber nitens M.F.Newman from Lao P.D.R. is described and illustrated.  
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L. Bai, J. Leong-Škorničková & N.H. Xia
Taxonomic studies on Zingiber (Zingiberaceae) in China I: Zingiber kerrii and the synonymy of Z. menghaiense and Z. stipitatum [Page 129 - 142]
A general introduction to Zingiber is provided, along with an outline of the materials and methods used in this paper and which are to be used in our subsequent taxonomic studies on Zingiber in China. Based on an investigation of the protologues, and on comparisons of both the type specimens and living materials collected from the type localities and their immediate vicinities, Zingiber stipitatum S.Q.Tong and Z. menghaiense S.Q.Tong are here reduced to synonymy under Z. kerrii Craib. A previous lectotypification of Zingiber kerrii is discussed and amended. The existence of mixed collections among the isotypes of Zingiber stipitatum is highlighted and a proposal to exclude the vegetative parts belonging to Z. neotruncatum from the type material is made. A description and a colour plate of Zingiber kerrii based on a collection from China are provided.
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C. Deori, N. Odyuo & A.A. Mao
Pennilabium labanyaeanum (Orchidaceae), a new species from Meghalaya, Northeast India [Page 143 - 149]
A new orchid species, Pennilabium labanyaeanum C.Deori, N.Odyuo & A.A.Mao, is described and illustrated from Meghalaya, India. A key to the species of Pennilabium in India is provided.
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K. Thoiba, A.K. Pradeep & C.N. Sunil
Tripogon bimucronatus (Poaceae: Chloridoideae: Tripogoninae), a new species from India [Page 151 - 157]
Tripogon bimucronatus Thoiba & Sunil, a new species of grass from the Nelliyampathy Hills, Palakkad District of Kerala, South India is described and illustrated. It is morphologically similar to Tripogon vellarianus Pradeep but differs by having adaxially scabrid, linear-acuminate leaves; ligules a fringe of hairs to 2.5 mm long; spikelets with 6–8 closely packed florets; lower glume asymmetrical; upper glume awned; lemma awn scabrid; palea elliptic-lanceolate, apex bimucronate and notched at the centre.
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K. Bransgrove & D.J. Middleton
A revision of Epithema (Gesneriaceae) [Page 159 - 229]
The genus Epithema Blume (Gesneriaceae) is revised. A key is given, all species are described and several are illustrated. Twenty species are recognised and two new combinations are made, Epithema philippinum (Hilliard & B.L.Burtt) Bransgrove and Epithema pusillum (C.B.)
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B.R. Maslin
Synoptic overview of Acacia sensu lato (Leguminosae: Mimosoideae) in East and Southeast Asia [Page 231 - 250]
Recent research shows that the formerly broadly circumscribed, pantropical genus Acacia Mill. (Leguminosae: Mimosoideae) is polyphyletic and should be treated as comprising at least five genera, namely, Acacia Mill. sensu stricto, Acaciella Britton & Rose, Mariosousa Seigler & Ebinger, Senegalia Raf. and Vachellia Wight & Arn. The indigenous flora of Acacia sensu lato in East and Southeast Asia comprise 52 species: 32 species (38 taxa) of Senegalia, 12 species of Acacia sensu stricto and eight species of Vachellia. These species are listed and their geographic distributions given, showing that Acacia sensu lato is unevenly distributed across the region, with centres of species-richness in Indonesia, Myanmar and Thailand. A summary of the classification history of Acacia sensu lato is provided and nomenclatural impacts of the recent retypification of Acacia briefly discussed.
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Jana Leong-Škorničková
BOOK REVIEW [Page 251 - 252]
Book Review:
Flowering Plants of the Western Ghats, Volume 1 (Dicots) T.S. Nayar, A. Rasiya Beegam & M. Sibi. 2014 & Flowering Plants of the Western Ghats, Volume 2 (Monocots). T.S. Nayar, M. Sibi & A. Rasiya Beegam. 2014.
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Year of Publication: 2014, Vol. 66 (2)

Date Published 20 November 2014
Y.F. Deng
Phlogacanthus magnus, a new combination in the Acanthaceae from Malaysia [Page 121 - 123]
An examination of the type material of Gymnostachyum magnum C.B.Clarke (Acanthaceae) from Malaysia reveals that it is a Phlogacanthus rather than a Gymnostachyum due to having two staminodes in addition to the two fertile stamens and muticous anther-thecae. The new combination, Phlogacanthus magnus (C.B.Clarke) Y.F.Deng is therefore proposed.
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R. Kiew
Two new species and one new subspecies of Ridleyandra (Gesneriaceae) from Peninsular Malaysia [Page 125 - 135]
Two new species, Ridleyandra bintangensis Kiew and R. nuangensis Kiew, from Kedah and Selangor respectively, and one new subspecies, R. kiewii subsp. magnifica Kiew, from Kelantan and Terengganu, are described from Peninsular Malaysia.

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R. Kiew
Two new white-flowered Codonoboea species (Gesneriaceae) from Peninsular Malaysia [Page 137 - 143]
Two new white-flowered Codonoboea species, C. mentigiensis Kiew and C. tembatensis Kiew (Gesneriaceae), are described from Peninsular Malaysia. Following the IUCN Criteria and Categories, C. tembatensis is Critically Endangered by clear-felling of forest prior to the area being flooded for an extension to a hydroelectric dam, while C. mentigiensis, that grows in lower montane forest within Permanent Protection Forest, is of Least Concern.

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J. Leong-Škorničková
A new lectotypification of Zingiber gracile var. elatius (Zingiberaceae) [Page 145 - 151]
The previous lectotypification of Zingiber gracile Jack. var. elatius Ridl. is deemed ineffective and a new lectotypification is here proposed. The original varietal epithet ‘elatior’, later also used at specific rank, is corrected here to ‘elatius’, to agree grammatically with the generic name.

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J. Leong-Škorničková, A. Thame & P.T. Chew
Notes on Singapore native Zingiberales I: A new species of Zingiber and notes on the identities of two further Zingiber taxa [Page 153 - 167]
A new species of Zingiber singapurense Škorničk., is described and illustrated here. It is compared to other species from the Zingiber gracile (Zingiber sect. Zingiber) alliance. A key to this group of species in Peninsular Malaysia and Singapore and a key to all Singapore native and naturalised Zingiber species are provided. The national and international conservation status of the new species and a conservation strategy for it are discussed. Notes on the varietal identity of Zingiber puberulum and the presence of Z. ottensii, previously misidentified as Z. zerumbet, in Singapore are given.
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S. Lindsay and C.W. Chen
Three new combinations in Haplopteris (Pteridaceae subfam. Vittarioideae) [Page 169 - 171]
Three new combinations are made here: Haplopteris alternans (Copel.) S.Linds. & C.W.Chen, Haplopteris humblotii (Hieron.) S.Linds. & C.W.Chen, and Haplopteris dareicarpa (Hook.) S.Linds. & C.W.Chen. Lectotypes for Vittaria alternans Copel. and Vittaria humblotii Hieron. are also designated here.

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R.O. Makinson and B.J. Conn
Puccinia psidii (Pucciniaceae – Eucalyptus Rust, Guava Rust, Myrtle Rust) – a threat to biodiversity in the Indo-Pacific region [Page 173 - 188]
A biotype of the South American fungal rust pathogen Puccinia psidii Winter has become widely naturalised along the east coast of Australia since early 2010, reaching North Queensland in 2012. This pathogen is known globally as Eucalyptus Rust or Guava Rust, and in Australia as Myrtle Rust. It is pathogenic on a wide range of plants in the Myrtaceae. This pathogen constitutes a major threat to myrtaceous plants in natural habitats and in production systems in moister areas of Australia, and potentially on a wide scale in the Malesian region. The risk of spread to the Malesian region, from Australia or other sources, is high, and once naturalised in Myrtaceae-rich biomes, eradication is unlikely.

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D.J. Middleton, J. Leong-Škorničková and Q.B. Nguyễn
A new species of Billolivia (Gesneriaceae) from Vietnam [Page 189 - 194]
The new species Billolivia moelleri D.J.Middleton from Núi Chúa National Park in Ninh Thuận Province in Vietnam is described. A provisional IUCN conservation assessment and a new key to the species of Billolivia are provided.

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M. Möller, W.-H. Chen, Y.-M. Shui, H. Atkins and D.J. Middleton
A new genus of Gesneriaceae in China and the transfer of Briggsia species to other genera [Page 195 - 205]
Since the transfer of several species of Briggsia to Oreocharis, including the type species Briggsia longifolia, the remaining 16 species of Briggsia have been in taxonomic limbo. We address this unsatisfactory situation by transferring 10 further species into Oreocharis on morphological grounds and by raising a new genus, with two species, based on previously published molecular data and their morphological distinction from other genera. This leaves only four species for which, at present, no satisfactory solution is available but, for pragmatic reasons until further research can be done, we place them in Loxostigma to which they are  morphologically most similar and in which one already has a combination. Several names are lectotypified.

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J.D. Mood, J.F. Veldkamp & L.M. Prince
A new species and a new record of Boesenbergia (Zingiberaceae) for Thailand [Page 207 - 214]
Boesenbergia purpureorubra Mood & L.M.Prince is described and illustrated. Boesenbergia longipes (King & Prain ex Ridl.) Schltr. is noted as a new record for Thailand based on C. Maknoi T38 (PSU, QBG).

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J.D. Mood, J.F. Veldkamp, S. Dey and L.M. Prince
Nomenclatural changes in Zingiberaceae: Caulokaempferia is a superfluous name for Monolophus and Jirawongsea is reduced to Boesenbergia [Page 215 - 231]
Wallich published Monolophus Wall. (Zingiberaceae) in 1832 for two taxa he had described previously in 1820 as Kaempferia linearis Wall. and K. secunda Wall. He also mentioned M. ? elegans Wall., but the “?” indicates that he was not certain that this species belonged to this new genus. Consequently, Monolophus elegans, while validly published, cannot be the lectotype of the generic name. In 1964 Larsen argued against accepting Monolophus and established the alternative Caulokaempferia K.Larsen. Caulokaempferia, typified by the name C. linearis (Wall.) K.Larsen, is, however, superfluous. Monolophus is hereby reinstated with 22 new combinations. Its phylogenetic position is shown in relation to other genera. Based on comparative nuclear and chloroplast DNA sequence data analyses, Jirawongsea Picheans. (previously Caulokaempferia, pro parte) is reduced to Boesenbergia Kuntze with five new combinations.

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L. Wijedasa, Z.Q. Shee and E. Chia
Conservation status and lectotypfication of Alangium ridleyi (Cornaceae) in Singapore [Page 233 - 239]
Alangium ridleyi King is lectotypified and the conservation status updated from Nationally Extinct to Endangered in Singapore and Endangered in Peninsula Malaysia.

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